| 100% | CITRON | 6 | Ancestor of the modern lemon and lime |
| 55% | EOHIPPUS | 8 | Ancestor of the modern horse |
| 52% | TARPAN | 6 | Wild ancestor of the modern-day horse |
| 51% | SHAWM | 5 | Medieval wind instrument; an ancestor of the modern oboe |
| 49% | VCR | 3 | Ancestor of the DVD player and DVR: Abbr. |
| 49% | VCR | 3 | Ancestor of the DVD player and TiVo: Abbr. |
| 46% | RINDS | 5 | Outsides of lemons and limes |
| 45% | UNCOLA | 6 | Geoffrey Holder described the lemon and lime he held in the 1970's 7 Up ads as these kind of "nuts" |
| 43% | ORANGES | 7 | __ and lemons, say the bells of St Clement's |
| 43% | ANDREALEVY | 10 | Author of the novels Fruit of the Lemon and Never Far from Nowhere |
| 41% | ADES | 4 | Lemon and lime drinks |
| 41% | ADES | 4 | Lemon and lime followers |
| 41% | LINNAEUS | 8 | Carolus —, 18th-century Swedish botanist who originated the modern scientific classification of plants and animals |
| 41% | ESAU | 4 | Ancestor of the Edomites |
| 41% | PANPIPE | 7 | Ancestor of the harmonica |
| 41% | LUTE | 4 | Ancestor of the guitar |
| 41% | GAUL | 4 | Ancestor of the French |
| 41% | NCAA | 4 | Org. at the center of modern "name, image and likeness" legislation |
| 41% | HUES | 4 | Lemon and lime, e.g. |
| 41% | LATIN | 5 | Ancestor of the romance languages |
| 40% | INCA | 4 | Ancestor of the Q'ero Indians |
| 40% | BASSHORN | 8 | Ancestor of the tuba or sousaphone |
| 40% | SOUR | 4 | Like lemons and limes |
| 39% | VIOL | 4 | Fretted ancestor of the fiddle |
| 39% | ENGLAND | 7 | Birthplace of the modern game of soccer (and winner of only one 24-Across) |
| 39% | GREECE | 6 | Birthplace of the modern Olympics |
| 39% | DDE | 3 | Inits. of the president whose ancestors were Pennsylvania Dutch and whose surname came from the German for "iron hewer" |
| 39% | SPRITE | 6 | Transparent lemon and lime carbonated drink |
| 39% | LGBTQ | 5 | Modern movement initialism ... or a hint to the starts of 18-, 24-, 40-, 52- and 61-Across |
| 39% | CIA | 3 | Org. that once funded the Museum of Modern Art, Partisan Review, and The Paris Review |